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We can observe that the only spot where the given sentence would fit is Blank 4. This sentence in question emphasises the broader, global significance of Central Asia's role; it reflects the idea that understanding the Silk Roads provides clarity on historical and global interconnectedness.
Since the given sentence shifts focus from describing the historical and cultural importance of the Silk Roads to their global implications, which is broader than the current discussion, placing it in any of Blanks 1, 2, or 3 interrupts the flow between the statements outlining certain features linked to the Silk Roads. For example, Blank 1 is a poor fit since the succeeding sentence mentions “These countries,” which refers to the “developing countries” mentioned in the sentence preceding the blank. Similarly, “They” in the sentence after Blank 2 refers to routes/Silk Roads described earlier. We can eliminate Blank 3 using a similar logic; the sentence after this blank establishes the metaphor of the Silk Roads as a central nervous system, emphasizing their global connectivity. The given sentence doesn’t align with the focus on the Silk Roads’ tangible impacts or the metaphorical depiction of their connectivity; therefore, placing it in Blank 3 would shift the focus prematurely to understanding their role globally.